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Subalpine Forest Composition Mapped by Imaging Spectroscopy

Combines hyperspectral remote sensing with field ecology to characterize the composition and structure of subalpine forests — aspen, lodgepole pine, and Engelmann spruce — across the Gunnison Basin, linking canopy properties to snow interception and watershed function.

Kebler PassCarbon PeakAlkali Basinimaging spectroscopyforest compositionutility valuePopulus tremuloidesPinus contortaPicea engelmanniiVegetation and Wildlife Studies for the Mount EmmoVegetation Appendix Materials for Vegetation and W1980Forest composition and structural controls on canoAgricultural Experiment Station

Knowledge Graph (283 nodes, 4978 connections)

Research Primer

Background

Subalpine forests in the Gunnison Basin of western Colorado form a mosaic of quaking aspen (Populus tremuloides), Engelmann spruce (Picea engelmannii), subalpine fir (Abies lasiocarpa), and lodgepole pine (Pinus contorta) that blanket elevations from roughly 2,700 to 3,500 meters. Managing these forests requires knowing not just where trees grow, but which species grow where, at what density, and in what condition. Imaging spectroscopy — high-fidelity remote sensing that captures hundreds of contiguous spectral bands using hyperspectral sensors — has emerged as a tool that allows land managers to map forest composition across entire watersheds at resolutions impossible with traditional field plots. Paired with concepts like self-thinning, root turnover, cover of bare ground, and sampling effort, spectroscopic maps let agencies estimate the utility value of stands for timber, wildlife habitat, watershed protection, and carbon storage.

This matters in the Gunnison Basin because forests here are under pressure from multiple directions at once: bark beetle disturbance is killing conifers in pulses that release needles and reshape biogeochemical cycles; beaver meadows and riparian corridors are shifting as hydrology changes; abandoned underground mines near Mount Emmons continue to influence soils and adjacent vegetation; and macroinvertebrate community composition in streams below forested slopes responds to changes in canopy cover and snowmelt timing. Mapping forest composition with spectroscopy — and validating those maps with ground frequency data on species, herbs, and shrubs — gives managers the baseline they need to track all of these changes.

Historical context

Systematic documentation of subalpine vegetation in the Gunnison Basin grew out of a combination of grazing studies, mining-related environmental assessments, and reclamation research. Early forage work such as the Determination of Annual Forage Production study on Nutras and Chavez Creeks (1954–1955) established baseline methods for quantifying ground cover composition, browse, and the effects of beaver habitat abandonment on montane meadows Determination of Annual Forage Production. In the late 1970s and early 1980s, the proposed molybdenum mine at Mount Emmons triggered some of the most detailed vegetation and wildlife inventories ever produced for the region, including the Vegetation and Wildlife Studies for the Mount Emmons Project (Keammerer & Stoecker, 1980) and its accompanying Vegetation Appendix with herb and shrub layer summaries for Mount Emmons, Alkali Basin, and Antelope Creek Basin Vegetation Appendix Materials.

Parallel efforts focused on how to restore high-elevation lands after disturbance. Technical work by Homestake Mining Company, Western State College, and the Colorado Mined Land Reclamation Board documented hydromulching, revegetation, and overburden management near Gunnison and Marshall Creek 1980 revegetation report, and AMAX-funded studies explored Potential Uses of Abandoned Underground Mines, including underground nursery facilities for high-altitude revegetation stock Potential Uses of Abandoned Underground Mines. Classification efforts culminated in A Classification of the Riparian Vegetation of the Gunnison River Basin (1995), produced with the Colorado Department of Natural Resources and the Environmental Protection Agency, which remains a foundational typology for riparian plant associations adjacent to subalpine forests Riparian Vegetation Classification.

Management actions and stakeholder roles

Forest composition management in the basin involves a layered set of actors. The U.S. Forest Service and the Colorado State Forest Service (CSFS) provide planting stock, reforestation guidance, and soil erosion control recommendations through publications such as Trees for Conservation: A Buyers Guide Trees for Conservation. The Rocky Mountain Biological Laboratory (RMBL) contributes long-term ecological monitoring, including targeted surveys such as the Small Mammal Survey at Acid Fen on Mt. Emmons, which linked habitat composition to vertebrate community structure Small Mammal Survey, Acid Fen. The Agricultural Experiment Station, Western State (now Western Colorado) University, and county and state land boards have historically supported applied research on forage, reclamation, and classification.

Management approaches combine remote sensing with field validation. Imaging spectroscopy products are now being calibrated against stand-level measurements of tree species identity, canopy structure, and understory composition — including common species like Thurber fescue (Festuca thurberi), silvery lupine (Lupinus argenteus), Wood rose (Rosa woodsii), and widespread non-natives such as Kentucky bluegrass (Poa pratensis) and dandelion (Taraxacum officinale). Riparian plant association keys Riparian Vegetation Classification and herb- and shrub-layer inventories from Mount Emmons Vegetation Appendix Materials serve as ground-truth references for spectral classification.

Current challenges and future directions

The most pressing challenges are climate-driven. Warming winters, earlier snowmelt, drought stress, and expanding bark beetle outbreaks are reshaping the relative dominance of aspen versus spruce-fir and lodgepole stands. Recent work at Kebler Pass using terrestrial laser scanning shows that canopy snow interception — a process that can return up to 50 percent of snowfall to the atmosphere via sublimation — differs sharply between deciduous aspen and coniferous species, meaning that shifts in forest composition directly alter watershed water yield (Beutler, 2024). Scaling those findings across the basin depends on accurate spectroscopic maps of which species occupy which slopes.

Future directions include integrating bark beetle disturbance tracking, post-fire regeneration monitoring, and legacy mining-impact assessments into a single compositional framework. Questions about self-thinning trajectories in dense lodgepole stands, root turnover in aspen clones, and expansion of bare ground after disturbance all require repeat spectroscopic surveys paired with plot-based ecology.

Connections to research

Imaging spectroscopy sits at the intersection of RMBL's long traditions in plant ecology, hydrology, and community monitoring. Spectral maps provide spatial context for RMBL's plot-based studies of subalpine meadows, aspen demography, and stream macroinvertebrate communities, while thesis research like Beutler (Beutler, 2024) demonstrates how composition controls the hydrologic function of mountain water towers. As agencies from the Forest Service to Colorado Parks and Wildlife adopt these data, the Gunnison Basin is becoming a testbed for linking hyperspectral forest inventories to decisions about water, wildlife, fire, and mine reclamation.

References

A Classification of the Riparian Vegetation of the Gunnison River Basin, Colorado.

Beutler, 2024. Forest composition and structural controls on canopy snow interception in a Colorado watershed.

Determination of Annual Forage Production.

Homestake/Western State revegetation and hydromulching technical report, 1980.

Potential Uses of Abandoned Underground Mines.

Small Mammal Survey, Acid Fen, Mt. Emmons, Gunnison, Colorado.

Trees for Conservation: A Buyers Guide.

Vegetation and Wildlife Studies for the Mount Emmons Project.

Vegetation Appendix Materials for Vegetation and Wildlife Studies for the Mount Emmons Project.

Species (187) →

Show 177 more speciess

Carex

sedges, sedgeCyperaceae · Poales · Plantae169 papers

Lupinus argenteus

silvery lupineFabaceae168 papers

Rosa woodsii

Wood Rose, Wood's roseRosaceae · Rosales · Plantae168 papers

Pascopyrum smithii

western wheatgrass, Western wheatgrassPoaceae · Poales · Plantae164 papers

Erythronium grandiflorum

avalanche lily, glacier lilyLiliaceae · Liliales · Plantae159 papers

Carex geyeri

elk sedgeCyperaceae · Poales · Plantae153 papers

Lonicera involucrata

Twinberry/Honeysuckle, bracted honeysuckleCaprifoliaceae · Dipsacales · Plantae153 papers

Lathyrus lanszwertii var. leucanthus

peavine, white-flowered peaFabaceae · Fabales · Plantae135 papers

Polemonium viscosum

Sky Pilot, alpine wildflowerPolemoniaceae · Ericales · Plantae133 papers

Wyethia amplexicaulis

mule's ears, Mules EarsAsteraceae · Asterales · Plantae130 papers

Aquilegia elegantula

red columbineRanunculaceae · Ranunculales · Plantae127 papers

Vaccinium myrtillus

grouse whortleberry, myrtle blueberryEricaceae · Ericales · Plantae124 papers

Picea pungens

Blue Spruce, Blue sprucePinaceae · Pinales · Plantae119 papers

Arnica cordifolia

heartleaf arnicaAsteraceae · Asterales · Plantae118 papers

Symphoricarpos oreophilus

mountain snowberryCaprifoliaceae · Dipsacales · Plantae118 papers

Ribes

gooseberry, CurrantGrossulariaceae · Saxifragales · Plantae118 papers

Trisetum spicatum

Spike triesetumPoaceae · Poales · Plantae114 papers

Veratrum californicum var. californicum

false hellebore, False HelleboreMelanthiaceae · Liliales · Plantae111 papers

Potentilla diversifolia

diverse-leaved cinquefoil110 papers

Thalictrum fendleri

Fendler meadowrueRanunculaceae · Ranunculales · Plantae107 papers

Rudbeckia montana

rayless, black-eyed susan, mountain coneflowerAsteraceae · Asterales · Plantae107 papers

Pseudotsuga menziesii

Douglas-fir, Douglas FirPinaceae · Pinales · Plantae104 papers

Festuca arizonica

Arizona fescuePoaceae · Poales · Plantae103 papers

Ligusticum porteri

lovage, host plantApiaceae · Apiales · Plantae103 papers

Nuphar polysepala

Waterlily, water lilyNymphaeaceae · Nymphaeales · Plantae102 papers

Prunus virginiana

ChokecherryRosaceae · Rosales · Plantae100 papers

Deschampsia cespitosa

Tufted Hairgrass, Tufted hairgrassPoaceae · Poales · Plantae99 papers

Achnatherum hymenoides

Indian Ricegrass, Indian ricegrassPoaceae · Poales · Plantae94 papers

Mertensia oblongifolia

green bluebellsBoraginaceae · Boraginales · Plantae93 papers

Poa fendleriana

Mutton grass, muttongrassPoaceae · Poales · Plantae89 papers

Pedicularis racemosa

parrots beak, curly-leaf lousewortOrobanchaceae · Lamiales · Plantae86 papers

Poa compressa

Canada Bluegrass, Canada bluegrassPoaceae · Poales · Plantae85 papers

Paxistima myrsinites

mountain loverCelastraceae · Celastrales · Plantae80 papers

Goodyera oblongifolia

rattlesnake plantainOrchidaceae · Asparagales · Plantae80 papers

Sambucus racemosa var. microbotrys

elderberryAdoxaceae · Dipsacales · Plantae80 papers

Lomatium dissectum

fern-leaf lomatiumApiaceae · Apiales · Plantae80 papers

Potentilla rubricaulis

Red-stem Cinquefoil, CinquefoilRosaceae · Rosales · Plantae80 papers

Agropyron scribneri

Plantae80 papers

Silene acaulis

Moss CampionCaryophyllaceae · Caryophyllales · Plantae80 papers

Saxifraga flagellaris

Whiplash SaxifrageSaxifragaceae · Plantae80 papers

Saxifraga cernua

Nodding SaxifrageSaxifragaceae · Saxifragales · Plantae80 papers

Minuartia rubella

Alpine SandwortCaryophyllaceae · Plantae80 papers

Minuartia obtusiloba

Twinflower SandwortCaryophyllaceae · Caryophyllales · Plantae80 papers

Selaginella densa

Plantae80 papers

Boechera oxylobula

RockcressBrassicaceae · Brassicales · Plantae80 papers

Arabis crandallii

Crandall RockcressCruciferae · Plantae80 papers

Achnatherum lettermanii

needlegrassPoaceae · Poales · Plantae80 papers

Achnatherum pinetorum

needlegrassPoaceae · Poales · Plantae80 papers

Rubus parviflorus

thimbleberryRosaceae · Rosales · Plantae80 papers

Festuca brachyphylla

alpine fescuePoaceae · Poales · Plantae80 papers

Geum rossii

alpine avensRosaceae · Rosales · Plantae80 papers

Aconitum columbianum

Monkshood, monkshoodRanunculaceae · Ranunculales · Plantae77 papers

Linum lewisii

Linaceae · Malpighiales · Plantae76 papers

Calamagrostis canadensis

Blue joint, Bluejoint reedgrassPoaceae · Poales · Plantae76 papers

Alnus tenuifolia

alder, narrow-leaf alder75 papers

Pinus ponderosa

ponderosa pine, Ponderosa PinePinaceae · Pinales · Plantae72 papers

Juniperus communis

Cupressaceae · Pinales · Plantae69 papers

Amelanchier alnifolia

ServiceberryRosaceae · Rosales · Plantae69 papers

Antennaria parvifolia

Asteraceae · Asterales · Plantae69 papers

Stipa

NeedlegrassPoaceae · Poales · Plantae67 papers

Helianthella quinquenervis

Asteraceae · Asterales · Plantae65 papers

Chamerion angustifolium ssp. angustifolium

Onagraceae · Myrtales · Plantae65 papers

Populus angustifolia

narrowleaf cottonwood, Narrowleaf CottonwoodSalicaceae · Malpighiales · Plantae64 papers

Marmota flaviventris

Yellow-bellied Marmots, yellow-bellied marmotSciuridae · Rodentia · Animalia64 papers

Eriogonum umbellatum

Sulpher-Flower, sulfur flower buckwheatPolygonaceae · Caryophyllales · Plantae60 papers

Carex aquatilis

water sedgeCyperaceae · Poales · Plantae59 papers

Artemisia frigida

Pasture Sagebrush, fringed sagebrushAsteraceae · Plantae57 papers

Aquilegia coerulea

Colorado Blue Columbine, ColumbineRanunculaceae · Ranunculales · Plantae57 papers

Trifolium pratense

Red Clover, red cloverFabaceae · Fabales · Plantae57 papers

Pseudocymopterus montanus

False Spring ParsleyApiaceae · Apiales · Plantae56 papers

Bromus tectorum

Cheatgrass, downy bromePoaceae · Poales · Plantae55 papers

Erigeron speciosus

Showy Fleabane, showy fleabaneAsteraceae · Asterales · Plantae55 papers

Carex sp.

sedgesPlantae54 papers

Zapus princeps

western jumping mice, Western Jumping MouseDipodidae · Rodentia · Animalia49 papers

Berberis repens

creeping barberryBerberidaceae · Ranunculales · Plantae48 papers

Castilleja linariifolia

Wyoming paintbrushOrobanchaceae · Lamiales · Plantae47 papers

Frasera speciosa

Monument plantGentianaceae · Gentianales · Plantae46 papers

Ranunculus inamoenus

ButtercupRanunculaceae · Ranunculales · Plantae46 papers

Penstemon strictus

Rocky Mountain penstemonPlantaginaceae · Lamiales · Plantae45 papers

Potentilla gracilis

graceful cinquefoilRosaceae · Rosales · Plantae44 papers

Galium circaezans var. circaezans

Rubiaceae · Gentianales · Plantae43 papers

Artemisia cana

silver sageAsteraceae · Asterales · Plantae43 papers

Fragaria virginiana ssp. glauca

Rosaceae · Rosales · Plantae43 papers

Antennaria rosea

Pussytoes, rosy pussytoesAsteraceae · Asterales · Plantae43 papers

Erigeron

Fleabane, fleabaneAsteraceae · Asterales · Plantae43 papers

Carex rossii

Plantae43 papers

Sedum lanceolatum

Stonecrop, yellow-flowered stonecrop plantCrassulaceae · Saxifragales · Plantae43 papers

Noccaea fendleri ssp. glauca

Brassicaceae · Brassicales · Plantae39 papers

Salix bebbiana

Bebb willow, bebb willowPlantae37 papers

Cynomys gunnisoni

Gunnison's prairie dogSciuridae · Rodentia · Animalia34 papers

Phenacomys intermedius

Silver, heather voleAnimalia34 papers

Microsorex hoyi

pigmy shrew, pygmy shrewAnimalia34 papers

Bromus ciliatus

Fringed Brome, fringed bromePoaceae · Poales · Plantae34 papers

Polemonium foliosissimum

leafy polemoniumPolemoniaceae · Ericales · Plantae33 papers

Salix planifolia

planeleaf willowSalicaceae · Malpighiales · Plantae31 papers

Equisetum arvense

Plantae31 papers

Phleum alpinum

Poaceae · Poales · Plantae31 papers

Viola canadensis

Violaceae · Malpighiales · Plantae31 papers

Pedicularis groenlandica

ElephantellaOrobanchaceae · Lamiales · Plantae31 papers

Thalictrum alpinum

alpine meadowrueRanunculaceae · Ranunculales · Plantae31 papers

Acer glabrum

Sapindaceae · Sapindales · Plantae31 papers

Osmorhiza depauperata

Apiaceae · Apiales · Plantae31 papers

Heracleum sphondylium ssp. montanum

cow parsnipApiaceae · Apiales · Plantae31 papers

Geranium richardsonii

Geraniaceae · Geraniales · Plantae31 papers

Conioselinum scopulorum

Plantae31 papers

Caltha leptosepala

western marsh marigoldRanunculaceae · Ranunculales · Plantae31 papers

Salix brachycarpa

barrenground willowSalicaceae · Malpighiales · Plantae31 papers

Bromus marginatus

Mountain bromePoaceae · Poales · Plantae31 papers

Geum triflorum

three-flowered avensRosaceae · Rosales · Plantae27 papers

Pinus flexilis

Limber Pine, limber pinePinaceae · Pinales · Plantae27 papers

Calochortus gunnisonii

Gunnison mariposa lilyLiliaceae · Liliales · Plantae27 papers

Hymenoxys hoopesii

Asteraceae · Asterales · Plantae27 papers

Bromus porteri

Poaceae · Poales · Plantae27 papers

Pedicularis grayi

Gray's LousewortOrobanchaceae · Lamiales · Plantae27 papers

Senecio bigelovii

Asteraceae · Asterales · Plantae27 papers

Viola adunca

Mountain Blue VioletViolaceae · Malpighiales · Plantae27 papers

Ribes inerme

CurrantGrossulariaceae · Saxifragales · Plantae26 papers

Sambucus racemosa

ElderberryAdoxaceae · Dipsacales · Plantae26 papers

Elymus trachycaulus

Slender wheatgrassPoaceae · Poales · Plantae26 papers

Senecio

Golden RagwortAsteraceae · Asterales · Plantae25 papers

Sialia currucoides

mountain bluebird, Mountain BluebirdTurdidae · Passeriformes · Animalia24 papers

Pinus edulis

Pinon Pine, Pinion PinePinaceae · Pinales · Plantae23 papers

Chamerion angustifolium

fire weed, fireweedOnagraceae · Myrtales · Plantae23 papers

Hordeum jubatum

Foxtail Barley, Foxtail barleyPoaceae · Poales · Plantae22 papers

Juniperus scopulorum

Eastern Red Cedar, Rocky Mountain juniperCupressaceae · Pinales · Plantae22 papers

Carex utriculata

beaked sedgeCyperaceae · Poales · Plantae22 papers

Pinus aristata

Bristlecone PinePinaceae · Pinales · Plantae21 papers

rose

rosePlantae21 papers

Mimulus guttatus

monkey flower, Common MonkeyflowerPhrymaceae · Lamiales · Plantae19 papers

Trifolium hybridum

Alsike Clover, Clover, alsikeFabaceae · Fabales · Plantae18 papers

Juniperus virginiana

eastern redcedar, Eastern Red CedarCupressaceae · Pinales · Plantae16 papers

Festuca idahoensis

Idaho fescuePoaceae · Poales · Plantae16 papers

Stipa comata

Needle and threadgrassPlantae16 papers

Stipa columbiana

Columbia needlegrassPlantae16 papers

Sitanion hystrix

Bottlebrush squirreltailPlantae16 papers

Populus balsamifera

poplarPlantae15 papers

Iris missouriensis

Wild Iris, irisIridaceae · Asparagales · Plantae14 papers

Caragana arborescens

Siberian Peashrub, CaraganaPlantae14 papers

Acer negundo

Box ElderSapindaceae · Sapindales · Plantae13 papers

Lagopus leucura

white-tailed ptarmigan, ptarmiganPhasianidae · Galliformes · Animalia13 papers

Erysimum capitatum

Brassicaceae · Brassicales · Plantae12 papers

Eleocharis palustris

creeping spikerushCyperaceae · Poales · Plantae12 papers

Salix boothii

Booth willowSalicaceae · Malpighiales · Plantae12 papers

Salix geyeriana

Geyer's willowSalicaceae · Malpighiales · Plantae12 papers

Wyethia arizonica

Arizona WyethiaAsteraceae · Asterales · Plantae12 papers

Thlaspi

PennycressBrassicaceae · Brassicales · Plantae11 papers

Elymus trachycaulus ssp. subsecundus

Slender Wheatgrass, Bearded wheatgrassPoaceae · Poales · Plantae11 papers

Zonotrichia leucophrys

white-crowned sparrow, Mountain White-crowned SparrowPasserellidae · Passeriformes · Animalia11 papers

Chrysothamnus viscidiflorus

green rabbitbrush, yellow rabbitbrushAsteraceae · Asterales · Plantae11 papers

Senecio integerrimus

GroundselAsteraceae · Asterales · Plantae11 papers

Muhlenbergia tricholepis

Hairy Dropseed, Pine dropseedPoaceae · Poales · Plantae11 papers

Claytonia lanceolata

Montiaceae · Caryophyllales · Plantae10 papers

Potentilla hippiana

woolly cinquefoil, ...Rosaceae · Rosales · Plantae10 papers

Rhus aromatica

SumacAnacardiaceae · Sapindales · Plantae10 papers

Bromus

brome grass, bromePoaceae · Poales · Plantae9 papers

Tetradymia canescens

gray horsebrushAsteraceae · Asterales · Plantae8 papers

Cirsium sp.

Thistle8 papers

Opuntia polyacantha

plains pricklypearCactaceae · Caryophyllales · Plantae8 papers

Elaeagnus angustifolia

Russian olive, Russian-oliveElaeagnaceae · Rosales · Plantae8 papers

Yucca glauca

small soapweed, Small SoapweedLiliaceae · Plantae8 papers

Phlox longifolia

long-leaf phlox, Long-leaved PhloxPolemoniaceae · Ericales · Plantae7 papers

Muhlenbergia richardsonis

mat muhly, Mat MuhlyGramineae · Plantae7 papers

Spinus pinus

pine siskin, Pine SiskinFringillidae · Passeriformes · Animalia7 papers

Ranunculus

ButtercupRanunculaceae · Ranunculales · Plantae6 papers

Betula occidentalis

water birchBetulaceae · Fagales · Plantae6 papers

Dasiphora fruticosa

Shrubby CinquefoilRosaceae · Rosales · Plantae6 papers

Senecio crassulus

Thickleaf GroundselAsteraceae · Asterales · Plantae5 papers

Box Elder

Box Elder, box elderPlantae5 papers

Bromus carinatus

Poaceae · Poales · Plantae5 papers

Solidago

goldenrod, GoldenrodAsteraceae · Asterales · Plantae5 papers

Lonicera tatarica

Honeysuckle, honeysucklePlantae4 papers

Syringa vulgaris

LilacOleaceae · Lamiales · Plantae4 papers

Spermophilus lateralis

Golden-mantled Ground Squirrel, golden-mantled ground squirrel4 papers

Myodes gapperi

red-backed vole, red-backed volesCricetidae · Rodentia · Animalia4 papers

Pooecetes gramineus

vesper sparrow, Vesper SparrowPasserellidae · Passeriformes · Animalia3 papers

Raphanus sativus

Radish, radishBrassicaceae · Brassicales · Plantae3 papers

Leymus cinereus

giant wildryePoaceae · Poales · Plantae3 papers

Stakeholder (1)

Agricultural Experiment Station

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